Spyro's other grandmother! Perle!

Perle, Selestina's mother, was a singer highly recognized and admired by water dragons. Although all water dragons sing and even form choirs, she stood out. She came to enchant Leviathan with her voice, whom she joined with. both singing each of their songs that they elaborated and dancing together.

A very kind and compassionate dragoness, she loved her daughter and mate Leviatán very much.

She died on the night of a thousand lamentations, saving her daughter from a water snake, being crushed to death by rocks to her and the water serpent.