Lito is based on a rat king (which is when a bunch of rats get stuck together by their tails). For some reason I always assumed it was just three rats, so I went with 3 limbs and tails as well as colors. It took a while to get the colors right so one wasn't overpowering the other. He's got a fun design though, I think one of my favs actually.
You can read more about him on Campfire